1. Taxonomy is the science of discovering, naming, describing, diagnosing, identifying, and classifying different kinds of taxa, from species to families. It lays the foundation for all of the biological sciences. The rapid increase in both taxonomic descriptions and malpractice in recent decades indicates a need for consistency in the procedure and quality of taxonomic research publications dealing with recognition of new taxa, name changes, and nomenclatural acts. Indeed, there are numerous examples in the recent literature of taxonomic works that fall short of the basic procedures and minimum standards required for naming new species according to the mandatory provisions and recommendations of the International Code of Zoological Nomenclature, accepted standards of taxonomic best practice, and journal editorial policy. Here, we provide practical guidelines of the procedure and key elements required to name, describe, and publish a new animal species or revise the taxonomic status.
  2. (Meyrick, 1909) and L. shikokuensis (Kawabe, 1984) in Taiwan. Two new species, Lepteucosma blandoides Lu & Hsu, sp. nov. and Lepteucosma dasyueshanensis Lu & Hsu, sp. nov., are found in Taiwan and being described. The key to the Lepteucsoma species in Taiwan is provided, and a checklist of the genus of the world is also given. We also propose a new combination, Lepteucosma yoshiyasui (Kawabe, 1989) comb. nov., based on the genitalia and wing characters. The results of this study increase the number of the Lepteucosma species to 24. Finally, we advocate paying more attention to the insects inhabitng the alpine area of Taiwan.
  3. Temperature is thought to be a key factor influencing global species richness patterns. We investigate the link between temperature and diversification in the butterfly family Pieridae by combining next generation DNA sequences and published molecular data with fine-grained distribution data. We sampled nearly 600 pierid butterfly species to infer the most comprehensive molecular phylogeny of the family and curated a distribution dataset of more than 800,000 occurrences. We found strong evidence that species in environments with more stable daily temperatures or cooler maximum temperatures in the warm seasons have higher speciation rates. Furthermore, speciation and extinction rates decreased in tandemwith global temperatures through geological time, resulting in a constant net diversification.
  4. Pachyrhabda citrinacma Meyrick, 1936, an endemic species to Formosa (Taiwan), was named by Edward Meyrick based on two specimens collected by Syuti Issiki. Despite some newly discovered stathmopodids exhibiting characteristics matching the original description, the lack of information regarding the type specimen depository led to the species remaining elusive since its initial naming. In this study, we successfully rediscovered a P. citrinacma specimen at the Insect Museum of National Taiwan University, where Syuti Issiki used to conduct research. The present article provides comprehensive morphological details for both sexes, as well as unveiling the genitalia characters and life history of this species for the first time. Larvae feed on spores of a range of fern species belonging to a few different families.
  5. The world’s largest butterfly genus Delias, commonly known as Jezebels, comprises ca. 251 species found throughout Asia, Australia, and Melanesia. Most species are endemic to islands in the Indo-Australian Archipelago or to New Guinea and nearby islands in Melanesia, and many species are restricted to montane habitats over 1200 m. We inferred an extensively sampled and well-supported molecular phylogeny of the group to better understand the spatial and temporal dimensions of its diversification. The remarkable diversity of Delias evolved in just ca. 15–16 Myr (crown age). The most recent common ancestor of a clade with most of the species dispersed out of New Guinea ca. 14 Mya, but at least six subsequently diverging lineages dispersed back to the island. Diversification was associated with frequent dispersal of lineages among the islands of the Indo-Australian Archipelago
  6. Apeculiar population of Ravenna nivea (Nire, 1920) was discovered from the Yinggeling Mountain Mass of central Hainan. Its wing pattern and COI barcode data show considerable distinction from other geographic populations of R. nivea, including that of Bawangling, approximately only 40 km away and also located in Hainan. The pdistance value of the COI barcode between the Yinggeling and Bawangling populations was 1.1%, considerably higher than the value (0.6%) between Bawangling population and populations in eastern China, where the subspecific name howarthi Saigusa, 1993 applies. The population is regarded as a distinct subspecies ngiunmoiae Lo & Hsu, subsp. nov. The distinctness and high degree of COI haplotype diversity of R. nivea found in Hainan and Taiwan suggest continental islands may serve as glacial refugees for the butterfly and other organisms during previous glaciations.
  7. Fern-spore-feeding (FSF) is rare and found in only four families of Lepidoptera. Stathmopodidae is the most speciose family that contains FSF species, and its subfamily Cuprininae exclusively specializes on FSF. However, three species of Stathmopodinae also specialize on FSF. To better understand the evolutionary history of FSF and, more generally, the significance of specialization on a peculiar host, a phylogenetic and taxonomic revision for this group is necessary. We reconstructed the most comprehensive molecular phylogeny, including one mitochondrial and four nuclear genes, of Stathmopodidae to date, including 137 samples representing 62 species, with a particular focus on the FSF subfamily, Cuprininae, including 33 species (41% of named species) from 6 of the 7 Cuprininae genera. Species from two other subfamilies, Stathmopodinae and Atkinsoniinae, were also included.
  8. Introduction of organisms to new range may impose detrimental effects on local organisms, especially when closely related species are involved. Species delimitation employing an integrative taxonomy approach may provide a quick assessment for the species status between taxa of interest, and to infer ecological competition and/or introgression that may be associated with the introduction. A nymphalid butterfly, Symbrenthia lilaea lunica, was recently introduced to Taiwan, where a closely related local taxon, S. l. formosanus, can be found.
  9. A new species of clearwing moth, Corematosetia parvimaculata sp. nov., is described from south-eastern China in this article. Adults and genitalia of both sexes are illustrated, DNA barcodes are provided, and immature biology and potential damage to Ardisia sieboldii Miq. and A. quinquegona Blume (Primulaceae) are both discussed.
  10. Butterflies are a diverse and charismatic insect group that are thought to have evolved with plants and dispersed throughout the world in response to key geological events. However, these hypotheses have not been extensively tested because a comprehensive phylogenetic framework and datasets for butterfly larval hosts and global distributions are lacking. We sequenced 391 genes from nearly 2,300 butterfly species, sampled from 90 countries and 28 specimen collections, to reconstruct a new phylogenomic tree of butterflies representing 92% of all genera. Our phylogeny has strong support for nearly all nodes and demonstrates that at least 36 butterfly tribes require reclassification. Divergence time analyses imply an origin ~100 million years ago for butterflies and indicate that all but one family were present before the K/Pg extinction event.
  11. Despite being the second largest group of vascular plants, ferns are scarcely reported being fed by insects when compared to angiosperms. Within these fern-feeding insects, lepidopterans are poorly represented and are restricted only to specific groups in this speciose order. The consumers specialising on fern spores are even scarcer in the order, with the majority being consumers of vegetative structures. Amongst the fern-spore-feeding Lepidoptera, Stathmopodidae is the family with the highest species diversity, even with a subfamily, Cyprininae Sinev, 2015, specialising on fern spores. However, fern-sporefeeding habit is not restricted to this subfamily. To understand the evolution of fern-sporefeeding within this family and to increase our knowledge of insect-fern evolution, detailed studies on fern-spore feeding stathmopodids are essential.
  12. Saridoscelis Meyrick, 1894 belongs to Yponomeutidae, with 9 described species. A new species, Saridoscelis diffusolinearum Lu & Hsu, sp. nov., is described from Taiwan, with diagnostic characters provided for both sexes herein. We also proposed that S. diffusolinearum sp. nov. is close related to S. kodamai Moriuti, 1961. The new species feeds on a subalpine bush Gaultheria cumingiana Vidal (Ericaceae) further indicating that the host plant of the subfamily Saridoscelinae may restricted to Ericaceae.
  13. An island famous for the work by a great pioneer naturalist Tadao Kano (鹿野忠雄),who suggested a “Neo-Wallace” line to run between Taiwan and Lanyu, has attracted many biologists to this small volcanic island. There are many names applied to the island. The aboriginal Tao people (called “Yami” by Ryuzo Torii【鳥居龍蔵】) calls it “Pongso no Tao”. Later 漢名 “ 紅頭嶼” were adopted. Another name “ 蘭嶼” was coined in 1947 for abundant orchids growing on the island, and “Lanyu” is the pronunciation of this name in Mandarin. It is known as “Botel Tobago” or “Orchid Island” in western countries. Moreover, there are a few additional names for Lanyu used by different aboriginal peoples in Taiwan. Consequently, Lanyu have many more “synonyms” than Taiwan itself.
  14. The year 2023 is an usually sad year to me, three persons important in my life has passed away: my best classmate friend as undergraduate student of National Taiwan University(臺灣大學),Mr. Benjamin Chen, my major professor in University of California, Dr. Jerry A. Powell and Mr. Kazuhiko Morishita, a teacher as well a longterm friend of mine (Fig. 1). My long lasting relationship with Mr. Morishita began when I was a college boy.
  15. Three newly recorded species of the genus Calicotis, Meyrick 1889 are reported from Taiwan: C. attiei (Guillermet, 2011), C. rotundinidus Terada, 2016, and C. exclamationis Terada, 2016. C. biserraticola Terada, 2016 is treated as a junior subjective synonym for C. attiei based on both morphological and molecular data. The life history of these three species is presented as well as the first observation of fern- feeding stathmopodid eggs in the world.
  16. Geographic and climatic differences between islands and continents may affect the evolution of their biota, and promote divergent selection in species distributed in both landscapes. To assess spatial-genetic structure, we genotyped 18 expressed sequence tag-simple sequence repeat (EST-SSR) loci and sequenced two mtDNA markers (ND5 and COI) and one nuclear marker (EF1α) in two subspecies of the butterfly Parantica sita.
  17. A new species of clearwing moth, Synanthedon suhua sp. nov., is described from Taiwan in this article. Adults and genitalia of both sexes are illustrated, DNA barcodes provided, and potential damage to Quercus longinux (Fagaceae) discussed.
  18. The immature stages, morphology, and systematic position of Bombyx incomposita (van Eecke, 1929) are reported. The morphology of larva was investigated using scanning electron microscopy, and the phylogeny was inferred using six genes (COI, CAD, EF-1α, GAPDH, RpS5, and wgl). The molecular phylogeny strongly supported that B. incomposita is sister to B. huttoni Westwood, 1847. The divergence of the COI barcode, phylogenetic reconstruction, and difference in immature stages support the current classification, in which the species incomposita belongs to the genus Bombyx. We give illustrations of male and female genitalia and briefly discuss the utility of the chaetotaxy of the first instar larvae on taxonomy of silkmoths. The SV group bears six setae in B. mandarina, but eight in B. incomposita. The L1 bears three setae in Rotunda, but just single seta in Bombyx.
  19. Deciduous Beech (Fagus spp.) forests have a highly disjunct distribution in the Northern Hemisphere, with greatest diversity in eastern Asia. Although Fagus supports a rich phytophagous insect fauna, the diversity, composition, and host specialization of insects associated with these trees in Asia are less well known compared with Europe and North America. For instance, two species of Catocala moths, both considered endemic to Japan, were previously known as the only beech-feeding specialists of this genus. However, two Catocala species were recently discovered from Taiwan during a survey of phytophagous insects on Taiwan Beech Fagus hayatae, a threatened plant. Identification of these two species of moths was determined by mtDNA (COI barcodes), adult morphology, including wing pattern and genitalia, and biology.
  20. Chrysozephyrus mushaellus paolongkoui, ssp. nov. is described and illustrated from Hainan island of southern China. This new subspecies possesses diagnosable characters different from previously recognized subspecies of C. mushaellus in wing marking and genitalia of both sexes. It utilizes Lithocarpus litseifolius (Fagaceae) as the larval hostplant.
  21. A taxonomic review of types of the clearwing moths described by Ji-kun Yang and Yin Wang from China is presented. Illustrations of types and their labels of 18 species are provided. For each species images of specimens and updated taxonomic information including the images and information on type specimens, complete bibliographies, data on host plants, distribution and current status are presented.
  22. between Taiwan and Japan, it is still unknown whether the nominate subspecies, P. s. sita, migrates seasonally in southwest China. To understand the potential migration of P. s. sita, it is necessary to investigate the genetic differentiation among populations. In our study, we developed 18 novel microsatellite markers from the transcriptome using next-generation sequencing methods.
  23. Two new species of Stathmopodidae are described from Taiwan: Cuprina atayalica Shen & Hsu, sp. nov., reared from larvae on Microsorum brachylepis, and C. insolita Hsu & Shen, sp. nov., reared from larvae on Tectaria subtriphylla and T. harlandii. Diagnostic characters for both species are provided. Larval host plants and the biology of the immature stages of both new species are documented.
  24. Three Graphium species belonging to two species groups of the subgenus Pazala, the alebion and tamerlanus groups, were examined in molecular and morphological studies, and their female genitalia are reported for the first time. Their relationship with other species groups within the subgenus is assessed and their divergence times are estimated. We find that G. (P.) alebion is the first lineage to diverge within Pazala in the early Miocene (20 Ma) and that G. (P.) tamerlanus and G. (P.) parus are sister species and diverged from each other in the late Miocene (7 Ma). A revision of the four recognised taxa belonging to
  25. Based upon both morphological and gene barcode evidence, the population of Spindasis syama in Taiwan is described as Spindasis syama lamuae, ssp. nov., and the taxon negrita Felder, 1862 is revised as a species endemic to the Philippines (stat. rev.).
  26. Lycaena nisa Wallace, 1866 was described from Formosa (Taiwan) and is here recognized as a senior subjective synonym of Lycaena alsulus Herrich-Schäffer, 1869. It is resurrected to serve as the valid name, Famegana nisa (Wallace, 1866), comb nov. of the species commonly known as Famegana alsulus. The name Zizera taiwana Sonan, 1938 (syn. nov.), also described from Formosa, is recognized as a junior subjective synonym of L. nisa. Another name, Zizeeria alsulus eggletoni Corbet, 1941 (syn. nov.), described from Hong Kong is also considered a junior subjective synonym of L. nisa. Moreover, all former synonyms of alsulus automatically become new junior synonyms for nisa. This species occurs in the Oriental and Australian regions and western Pacific.
  27. Fifteen holotypes of Asian Hesperiidae taxa described by Shu-iti Murayama were examined, and their taxonomic status is discussed. We confirm that five are valid names while the rest are synonyms as indicated partially by previous authors. Valid species-level names are Aeromachus matudai (Murayama), Aeromachus bandaishanus Murayama & Shimonoya, Coladenia pinsbukana (Shimonoya & Murayama), and Sebastonyma suthepiana Murayama & Kimura. Valid subspecieslevel name is Ochlodes yuchingkina Murayama & Shimonoya. In order to settle the taxonomic status of Pedesta masuriensis cuneomaculata Murayama, masuriensis and tali were studied morphologically and molecularly. As a result, we consider that masuriensis and tali are two different species and treat cuneomaculata as a junior subjective synonym of tali.
  28. The genus Epsteinius Lin, Braby & Hsu gen. nov. is established to accommodate a new, diminutive species of limacodid moth, E. translucidus Lin, sp. nov., from central Taiwan. Although the new species is the sister taxon to Microleon Butler, 1885 according to molecular phylogenetic data, it differs fundamentally from that genus in adult morphology, wing color pattern, male and female genitalia, and larval morphology. Larvae of E. translucidus Lin, sp. nov. represent an interesting case in which crypsis is based on transparency through its body color, resting behavior, and feeding pattern.
  29. Ussuriana fani wangi, ssp. nov. is described and illustrated from Baihuashan of Taihangshan Mountain Range, Beijing. This new subspecies utilizes Fraxinus bungeana as the larval hostplant, representing the first record of U. fani in northern China. The immature biology, larval morphology, and data of COI barcode all indicate the population belonging to U. fani rather than superficially similar U. michaelis.
  30. Neptis reducta Fruhsorfer, 1908 is currently recognized as a nymphalid butterfly found only in Taiwan and Thailand, with previously recorded larval hostplant and immature biology involving mis-identifications. The present study recognizes Aphananthe aspera (Cannabaceae) as the sole larval hostplant for N. reducta, and the nymphalid butterfly is the first known lepidopterous insect specialized on Aphananthe.
  31. Araragi panda sichuanensis Hsu & Li, subsp. nov. (Theclinae: Theclini) is described from Baoxing, Sichuan Province of western China, the type locality of the Giant Panda Ailuropoda melanoleuca. The present article also clarifies the identity of the hostplant for A. panda previously mis-identified. It should be the wheel wingnut (Cyclocarya paliurus) instead of the iron walnut (Juglans sigllata) recorded in literature. C. paliurus is sometimes called “the Giant Panda in plant kingdom” in China.
  32. Ants are a dominant insect group in terrestrial ecosystems and many myrmecophilous species evolve to associate with ants to gain benefits. One iconic example is myrmecophilous butterflies that often produce ant-mimicking vibrational calls to modulate ant behaviors. Despite its popularity, empirical exploration of how butterflies utilize vibrational signals to communicate with ants is scarce. In this study, we reported that the myrmecophilous butterfly Spindasis lohita produce three types of larval calls and one type of pupal call, while its tending ant, Crematogaster rogenhoferi emit a single type of call. The results of discriminant analysis revealed that calls of the two species are quantitatively similar in their signal attributes; the potential role of butterfly calls are further confirmed by the playback experiments in which certain ant behaviors including antennation, aggregation, and guarding were induced when one of the butterfly calls was played to C. rogenhoferi worker.
  33. Larvae of the cosmopolitan family Limacodidae, commonly known as “slug” caterpillars, are well known because of the widespread occurrence of spines with urticating properties, a morpho‐chemical adaptive trait that has been demonstrated to protect the larvae from natural enemies. However, while most species are armed with rows of spines (“nettle” caterpillars), slug caterpillars are morphologically diverse with some species lacking spines and thus are nonstinging. It has been demonstrated that the evolution of spines in slug caterpillars may have a single origin and that this trait is possibly derived from nonstinging slug caterpillars, but these conclusions were based on limited sampling of mainly New World taxa; thus, the evolution of spines and other traits within the family remains unresolved.
  34. Taiwan is at the north-eastern limits of the Asian center of diversity for fruit flies in the tribe Dacini, and the country has several endemic species. Between 2013–2015, we surveyed the diversity of dacine fruit flies in Taiwan using kairomone baited traps and found fifteen species. We report four species for the first time in Taiwan and figure them: Bactrocera nigrifacia Zhang, Ji & Chen 2011, B. rubigina (Wang & Zhao 1989), B. dorsaloides (Hardy & Adachi 1954) and B. bhutaniae Drew & Romig 2013. For the species that are classified as pests, we explored how their spatial distribution correlates with elevation. The oriental fruit fly, Bactrocera dorsalis (Hendel 1912), is the most abundant pest on the island, but decreases sharply in abundance at higher elevations. Other pest species occur in much smaller numbers and respond differently to elevation and latitude.
  35. Caltoris ranrunna (Sonan, 1936) is resurrected from synonymy of Caltoris cahira austeni (Moore, 1883) to represent a Caltoris species endemic to Taiwan based upon COI barcode divergence and morphological diagnosis in genitalia of both sexes. This species distributes allopatrically from C. cahira of continental Asia and Andamans.
  36. Two new species of the tribe Synanthedonini are described from Taiwan: Synanthedon auritinctaoidis sp.
  37. The life history, morphology, and biology of the immature stages and phylogenetic relationships of Rotunda rotundapex (Miyata & Kishida, 1990) are described and illustrated for the first time. The species is univoltine: eggs hatch in spring (March or April) and the life cycle from egg to adult is completed in about 3 wk, with larvae developing rapidly on young leaves of the host plants, Morus australis and to a lesser extent Broussonetia monoica (Moraceae), and adults emerging in April–May. Eggs are laid in clusters on twigs of the host plant, are covered by scales during female oviposition, and remain in diapause for the remainder of the year (i.e., for 10–11 mo).
  38. Mycalesis kagina Fruhstorfer, 1911 is separated from Mycalesis suaveolens Wood-Mason & de Nicéville, 1883 to represent a species endemic to Taiwan based upon COI barcode divergence, morphological diagnosis of larva, and genitalia of both sexes. Both kagina and suaveolens are confirmed as members of the genus Mycalesis in Mycalesina. Immature morphology, biology, and hostplant associations for both species are given for the first time. Larvae of both species are recognized as specialists on Zingiberaceae, a plant family rarely used by satyrid butterflies.
  39. We reared a Telenomus species from eggs of Bombyx mandarina (Moore) (Lepidoptera: Bombycidae) and Bombyx mori (Linnaeus) (Lepidoptera: Bombycidae) in Japan, and from eggs of B. mandarina in Taiwan. Morphological examination revealed that this Telenomus species is new to science. In this article, we describe it as Telenomus moricolus Matsuo et Hirose, sp. nov. Because B. mandarina is considered to be an ancestor of B. mori, a domestic insect, it is reasonable to assume that B. mandarina is an original host of T. moricolus. This is the second discovery of an egg parasitoid attacking wild and domesticated silkworms, following the first discovery of T. theophilae, a Chinese species. The significance of the discovery of T. moricolus is discussed in relation to examining the effects of host-insect domestication on egg parasitism.
  40. - Li, A., H. Deng, Y. F. Hsu. 2018. Discovery of a Crateva-associated colony of Pieris brassicae (Linnaeus, 1758) from southwestern Yunnan (Lepidoptera: Pieridae). Butterflies 76: 52-55.
  41. - Hsu, Y. F. A New Species of Chrysozephyrus associated with Lithocarpus corneus (Lepidoptera: Lycaenidae: Theclini). Zootaxa 4377 (1): 143–150.
  42. - Wu, L. W., W. J. Lin, Y. F. Hsu*. 2018. A distinct species, Dodona formosana, detected in the Dodona eugenes species complex: clarification of the taxonomic status of the Punch butterfly in Taiwan. ZooKeys 736: 59–77.
  43. Liang, J. Y., Y.F. Hsu. Two new species of clearwing moths (Lepidoptera: Sesiidae) from Taiwan. Zootaxa 4299 (3): 415–422.
  44. Chen, Y. F., C. L. Huang, Y. F. Hsu. DNA barcoding and morphological data reveal a new Hyposoter (Hymenoptera: Ichneumonidae: Porizontinae) reared from a rare zygaenid moth Artona flavipuncta Hampson, 1900 in Taiwan. Zootaxa 4337 (2): 279–287
  45. Chen, Y. F., C. L. Huang, Y. F. Hsu. DNA barcoding and morphological data reveal a new Hyposoter (Hymenoptera: Ichneumonidae: Porizontinae) reared from a rare zygaenid moth Artona flavipuncta Hampson, 1900 in Taiwan. Zootaxa 4337 (2): 279–287
  46. Hsu, Y. F., H. C. Huang, C. C. Lu. 2017. On mysterious occurrence of butterflies in the genus Delias at Jinmen Islands, with clarification on the nomenclature of D. pasithoe (Linnaeus, 1767) and D. acalis (Godart, 1819) in southern China. Zootaxa 4276 (2): 245–254.
  47. Wu, S, WC Chang, LH Wang, CL Huang, YF Hsu*. 2016. Descriptions of two new notodontid species from the relic Fagus forests in northeastern Taiwan (Lepidoptera). Zootaxa 4066(3): 291–300.
  48. Arita, Y., A. Kallies, Y. F. Hsu, J. Y. Liang, B. C. Lai, M. M. Yang, N. Yata. Polymorphism of Nokona pilamicola (Strand, [1916]) (Lepidoptera, Sesiidae) in Taiwan. Tinea 23 (5): 249-254.
  49. Hsu YF, CL Huang, HC Huang. 2016. Immature biology and morphology of an obligate myrmecophilous butterfly Catapaecilma major moltrechti (Wileman) (Lepidoptera: Lycaenidae) from Taiwan. Zootaxa 4139 (2): 274–284.
  50. Huang, C. L, Y. F. Hsu*. 2016. Immature biology of Neptis hesione podarces (Lepidoptera: Nymphalidae) in Taiwan, with discussion on its frass chain function. Annals of Entomological Society of America. 2016 saw 010: 1-9.
  51. Liang, J. Y. & Y. F. Hsu*. 2015. A review of clearwing moths in the tribe Synanthedonini, with descriptions of six new species from Taiwan (Lepidoptera: Sesiidae). Zootaxa 4044(4): 535–555.
  52. Hsu, Y. F. 2015. A new species of Shaanxiana (Lepidoptera: Lycaenidae) from southern China. Zootaxa 4027(1): 130-134.
  53. Espeland, M., J. P.W. Hall, P. J. DeVries, D. C. Lees, M. Cornwall, Y. F. Hsu, L.W. Wu, D. L. Campbell, G. Talavera, R. Vila, S. Salzman, S. Ruehr, D. J. Lohman, N. E. Pierce. 2015. Ancient Neotropical origin and recent recolonisation: Phylogeny, biogeography and diversification of the Riodinidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 93 (2015) 296–306. doi:10.1016/j.ympev.2015.08.006
  54. Wu, L. W., S. H. Yen, D. C. Lees, C. C. Lu, P. S. Yang, Y. F. Hsu*. 2015. Phylogeny and historical biogeography of Asian Pterourus butterflies (Lepidoptera: Papilionidae): a case of intercontinental dispersal from North America to East Asia. PLoS ONE 10(10): e0140933. doi:10.1371/journal.pone.0140933
  55. Wu, L. W., L. H. Lin, D. C Lees, Y. F. Hsu. Mitogenomic sequences effectively recover relationships within brush-footed butterflies (Lepidoptera: Nymphalidae) BMC Genomics 2014, 15:468
  56. Tan, W. H., T. H. Liu, Y. K. Lin, Y. F. Hsu. 2014. Restoration of an endangered plant, Hygrophila pogonocalyx, leads toan adaptive host shift of the chocolate pansy (Junonia iphita iphita). Zoology 117: 237-244.
  57. 千葉秀幸、徐堉峰、黃行七。蝴蝶王囯台湾の蛻変。昆虫と自然 49(14): 22-31.
  58. Hsu, Y. F., C. K. Wang, C. L. Huang. Notes on Ypthima lisandra from Kinmen Islands (Lepidoptera: Nymphalidae: Satyrinae). Butterflies (S. fujisanus) 63: 49– 53.
  59. Hsu, Y. F., H. C. Huang. 2013. Prosotas gracilis gracilis Röber, 1886, a new record of blue butterfly found in Lanyu, Taiwan (Lepidoptera, Lycaenidae, Lycaeninae, Polyommatini). BioFormosa 48: 13-16.
  60. Hsu, Y. F., J. H. Lin1, I. H. Wu, P. S. Yang. 2013. Mycalesis igoleta C. & R. Felder, the first satyrid vagrant butterfly visiting Lanyu, Taiwan (Lepidoptera: Nymphalidae: Satyrinae) Butterflies (Teinopalpus) 64: 33–35.
  61. Hsu, Y. F., H. C. Huang. 2013. Rapala manea schistacea (Moore, 1879), a hairstreak butterfly new to Kinmen Islands (Lycaenidae: Theclinae): Butterflies (Teinopalpus) 61: 40-42.
  62. Daimon T, T. Fujii, M. Yago, Y. F. Hsu, Nakajima, T. Fujii, S. Katsuma, Y. Ishikawa, T. Shimada. 2012. Female sex pheromone and male behavioral responses of the bombycid moth Trilocha varians: comparison with those of the domesticated silkmoth Bombyx mori. Naturwissenschaften 99(3): 207–215. doi: 10.1007/s00114-012-0887-3
  63. Daimon T, M Yago, YF Hsu, T Fujii, Y Nakajima, R Kokusho, H Abe, S Katsuma, T Shimada. 2012. Molecular phylogeny, laboratory rearing, and karyotype of the bombycid moth, Trilocha varians (Lepidoptera: Bombycidae: Bombycinae). Journal of Insect Science 12: 1-17. doi: 10.1673/031.012.4901
  64. Huang, C. L. & Y. F. Hsu*. 2011. The early stages and biology of Minois nagasawae (Matsumura)(Lepidoptera: Nymphalidae, Satyrinae), an alpine butterfly endemic to Taiwan. Proceedings of the Entomological Society of Washington 113: 325-334. DOI: 10.4289/0013-8797.113.3.325
  65. Wu, L. W., D. C. Lees, S. H. Yen, C. C. Lu & Y. F. Hsu*. 2011. The complete mitochondrial genome of the endangered swallowtail, Agehana maraho (Lepidoptera: Papilionidae): evaluating sequence variability and suitable markers for conservation genetic studies. Entomological News 121: 267-280. doi: http://dx.doi.org/10.3157/021.121.0308
  66. Wu, L. W., S. H. Yen, D. C. Lees & Y. F. Hsu*. 2010. Elucidating genetic signatures of native and introduced populations of the Cycad Blue, Chilades pandava to Taiwan: a threat both to Sago Palm and to native Cycas populations worldwide. Biological Invasions 12: 2649-2669.
  67. Wu, L. W., S. H. Yen, D. C. Lees & Y. F. Hsu*. 2010. Elucidating genetic signatures of native and introduced populations of the Cycad Blue, Chilades pandava to Taiwan: a threat both to Sago Palm and to native Cycas populations worldwide. Biological Invasions 12: 2649-2669.
  68. Hsu, Y. F. 2009. A new species of Antigius (Lepidoptera: Lycaenidae: Theclini) from Taiwan. Zootaxa 1983: 45-53.
  69. Chiba, H., Y. F. Hsu*, H. Tsukiyama, P. Y.-F. Lo, C. R. Chen, S. M. Wang. 2009. Jinhaku Sonan’s Skipper Type Collection Deposited at Taiwan. Agricultural Research Institute (Lepidoptera: Hesperiidae). Zootaxa 2202: 48-58.
  70. Hsu, Y. F.* & H. C. Huang. 2008. On the discovery of Hasora mixta limata ssp. nov. from Lanyu, Taiwan, with Observations on its unusual immature biology (Lepidoptera: Hesperiidae: Coeliadinae). Zoological Studies 47: 222-231.
  71. Hsu, Y. F.*, S. C. Chen & C. C. Lu. 2007. On the immature biology of Pseudocoladenia dan sadakoe (Sonan & Mitono), with a discussion of the systematic status of this rare taxon (Lepidoptera: Hesperiidae). Pan-Pacific Entomologist 83: 19-31.
  72. Hsu, Y. F. & S. H. Yen*. 2006. Immature biology of Prosotas reveals an oviposting strategy unusual in polyommatine lycaenids (Lepidoptera, Lycaenidae,Polyommatinae), with description of a new subspecies of P. dubiosa from Taiwan. Zoological Studies 45: 308-321.
  73. Hsu, Y. F.*, H. Chiba & S. H. Yen. 2006. Case 3339 Celaenorrhinus ratna Fruhstorfer, 1908 (Insecta, Lepidoptera): proposed conservation of the specific name. Bulletin of Zoological Nomenclature 63(2): 114-117.
  74. Hsu, Y. F., H. Chiba, Y. C. Yang, H. Tsukiyama, P. S. Yang & S. M. Wang. 2006. A Taxonomic Review of the Taiwanese Skippers Described by Shonen Matsumura (Lepidoptera: Hesperiidae). Zoological Studies 45: 322-338.

名稱 開始日期 結束日期 委託單位 編號 參與身份別
重要經濟昆蟲蠶蛾及其近緣物種之起源與演化(2/3) 2019/08/01 2020/07/31 科技部(原國科會) 107-2313-B-003 -002 -MY3 主持人
重要經濟昆蟲蠶蛾及其近緣物種之起源與演化(1/3) 2018/08/01 2019/07/31 科技部(原國科會) 107-2313-B-003 -002 -MY3 主持人
農林害蟲透翅蛾之多樣性與系統分類(3/3) 2018/08/01 2019/07/31 科技部(原國科會) 105-2313-B-003 -001 -MY3 主持人
農林害蟲透翅蛾之多樣性與系統分類(2/3) 2017/08/01 2018/07/31 科技部(原國科會) 105-2313-B-003 -001 -MY3 主持人
農林害蟲透翅蛾之多樣性與系統分類(1/3) 2016/08/01 2017/07/31 科技部(原國科會) 105-2313-B-003 -001 -MY3 主持人
東喜馬拉雅與台灣生物區系隔離分化的式樣與形成機制-基於代表昆蟲(兩岸合作研究 2015/01/01 2018/12/31 科技部(原國科會) 104-2621-B-003-001-MY3 主持人
以寬尾鳳蝶屬的親緣關係及親緣地理學探討洲際間的隔離/散佈形況(II)(2/2) 2014/08/01 2015/07/31 科技部(原國科會) 102-2621-B-003-004-MY2 主持人
建構社會生活場域中的生物監測機制—ICCA在蘭嶼的長期生物監測(II) 2013/08/01 2014/07/31 科技部(原國科會) 102-2621-M-003-003- 主持人
以寬尾鳳蝶屬的親緣關係及親緣地理學探討洲際間的隔離/散佈形況(II)(1/2) 2013/08/01 2014/07/31 科技部(原國科會) 102-2621-B-003-004-MY2 主持人
建構社會生活場域中的生物監測機制—ICCA在蘭嶼的長期生物監測(I) 2012/08/01 2013/07/31 科技部(原國科會) 101-2621-M-003-004- 主持人
以寬尾鳳蝶屬的親緣關係及親緣地理學探討洲際間的隔離/散佈形況 2012/08/01 2013/07/31 科技部(原國科會) 101-2621-B-003-001- 主持人
以形態及分子證據探討翠灰蝶族之系統演化與系統分類(3/3) 2010/08/01 2011/07/31 科技部(原國科會) 97-2621-B-003-003-MY3 (97B0282-03) 主持人
臺灣蝶類資源之分類系統修訂、分類地位釐清與名錄整理﹝二﹞ 2010/01/01 2010/12/31 其他政府部門 99林管-02.1-保-23(5) 主持人
以形態及分子證據探討翠灰蝶族之系統演化與系統分類(2/3) 2009/08/01 2010/07/31 科技部(原國科會) 97-2621-B-003-003-MY3 (97B0282-02) 主持人
臺灣蝶類資源之分類系統修訂、分類地位釐清與名錄整理﹝一﹞ 2009/01/01 2009/12/31 其他政府部門 98林管-02.1-保-19(9) 主持人
以形態及分子證據探討翠灰蝶族之系統演化與系統分類(1/3) 2008/08/01 2009/07/31 科技部(原國科會) 97-2621-B-003-003-MY3(97B0282-01) 主持人
台灣野生物遺傳多樣性與保育遺傳研究:台灣產翠蛺蝶族遺傳多樣性與親緣關係之研究(III) 2008/01/01 2008/12/31 其他政府部門 97農科-11.1.2-務-e1 總主持人
台灣陸域之節肢動物之生物多樣性及生物地理學研究-利用分子,形態及食性之綜合證據探討疑問種黃蝶之多樣性問題及生物地理研究(3/3) 2007/08/01 2008/07/31 科技部(原國科會) 96-2621-B-003-001- 主持人
台灣野生物遺傳多樣性與保育遺傳研究:台灣產翠蛺蝶族遺傳多樣性與親緣關係之研究(II) 2007/01/01 2007/12/31 其他政府部門 96農科-11.1.2-務-e1(3) 主持人
密紋波眼蝶種群之系統關係重建及分類問題 2006/08/01 2007/07/31 科技部(原國科會) 95-2621-B-003-001- 主持人
台灣陸域之節肢動物之生物多樣性及生物地理學研究-利用分子,形態及食性之綜合證據探討疑問種黃蝶之多樣性問題及生物地理研究(2/3) 2006/08/01 2007/07/31 科技部(原國科會) 95-2621-B-003-002- 主持人
台灣野生物遺傳多樣性與保育遺傳研究:台灣產翠蛺蝶族遺傳多樣性與親緣關係之研究(I) 2006/01/01 2006/12/31 其他政府部門 95農科-11.1.3-務-e1(4) 主持人
密紋波眼蝶種群之系統關係重建及分類問題(2/3) 2005/08/01 2016/07/31 科技部(原國科會) 94-2621-B-003-002- 主持人
台灣陸域之節肢動物之生物多樣性及生物地理學研究-利用分子,形態及食性之綜合證據探討疑問種黃蝶之多樣性問題及生物地理研究(1/3) 2005/08/01 2006/07/31 科技部(原國科會) 94-2621-B-003-004- 主持人
密紋波眼蝶種群之系統關係重建及分類問題(1/3) 2004/08/01 2005/07/31 科技部(原國科會) 93-2621-B-003-003- 主持人
熱帶/亞熱帶地區之關聯族群研究:蘇鐵小灰蝶族群結構與動態(3/3) 2003/08/01 2004/07/31 科技部(原國科會) 92-2313-B-003-001- 主持人
熱帶/亞熱帶地區之關聯族群研究:蘇鐵小灰蝶族群結構與動態(2/3) 2002/08/01 2003/07/31 科技部(原國科會) 91-2313-B-003-002- 主持人
熱帶/亞熱帶地區之關聯族群研究:蘇鐵小灰蝶族群結構與動態(1/3) 2001/08/01 2002/07/31 科技部(原國科會) 90-2313-B-003-002- 主持人
植物與昆蟲交互作用之系統發生研究: 水青岡與小灰蝶之協同進化(三) 2000/08/01 2001/07/31 科技部(原國科會) 89-2311-B-003-019- 主持人
植物與昆蟲交互作用之系統發生研究: 水青岡與小灰蝶之協同進化(二)) 1999/08/01 2000/07/31 科技部(原國科會) 89-2311-B-003-008- 主持人
植物與昆蟲交互作用之系統發生研究: 水青岡與小灰蝶之協同進化 1998/08/01 1999/07/31 科技部(原國科會) 88-2311-B-003-010- 主持人
台灣陸生生物進化之探討--莫氏波紋蛇目蝶種群之親緣關係與族群變異之研究(II) 1997/08/01 1998/07/31 科技部(原國科會) 87-2311-B-003-016-B17 主持人
台灣地區琉璃鳳紋蝶異質族群之演化及分類討論 1996/08/01 1997/07/31 科技部(原國科會) 86-2311-B-018-003- 主持人
台灣陸生生物進化之探討--莫氏波紋蛇目蝶種群之親緣關係與族群變異之研究 1996/08/01 1997/07/31 科技部(原國科會) 86-2311-B-018-001-B17 主持人

  1. 徐堉峰,梁家源,黃智偉,沈宗諭。2021。《臺灣蝶類誌第五卷:蛺蝶科》內容均依正統分類學方式製作,按科屬種方式編排,每一蝶種介紹,包含中名、異名表列、標本檢查、成蟲形態、世界分佈、在台灣分布、幼蟲寄主植物、生物學及註記;而且每一種類都有雌雄背、腹面標本彩圖、交尾器解剖彩圖,是首部完整論述臺灣本島與離島蝶類資源的分類學專著,將分年編纂與出版,中英對照,共計五卷。
  2. 徐堉峰,梁家源,黃智偉。2020。《臺灣蝶類誌第四卷:灰蝶科》內容均依正統分類學方式製作,按科屬種方式編排,每一蝶種介紹,包含中名、異名表列、標本檢查、成蟲形態、世界分佈、在台灣分布、幼蟲寄主植物、生物學及註記;而且每一種類都有雌雄背、腹面標本彩圖、交尾器解剖彩圖,是首部完整論述臺灣本島與離島蝶類資源的分類學專著,將分年編纂與出版,中英對照,共計五卷。
  3. 徐堉峰,千葉秀幸,築山 洋,梁家源,黃智偉。2019。《臺灣蝶類誌第三卷:蛺蝶科》內容均依正統分類學方式製作,按科屬種方式編排,每一蝶種介紹,包含中名、異名表列、標本檢查、成蟲形態、世界分佈、在台灣分布、幼蟲寄主植物、生物學及註記;而且每一種類都有雌雄背、腹面標本彩圖、交尾器解剖彩圖,是首部完整論述臺灣本島與離島蝶類資源的分類學專著,將分年編纂與出版,中英對照,共計五卷。
  4. 徐堉峰、許育銘。陽明飛蛾。2017。陽明山國家公園管理處。320頁。ISBN 9789860550801,GPN 1010602830。
  5. 徐堉峰,黃嘉龍,梁家源。2018。《臺灣蝶類誌第一卷:鳳蝶科》內容均依正統分類學方式製作,按科屬種方式編排,每一蝶種介紹,包含中名、異名表列、標本檢查、成蟲形態、世界分佈、在台灣分布、幼蟲寄主植物、生物學及註記;而且每一種類都有雌雄背、腹面標本彩圖、交尾器解剖彩圖,是首部完整論述臺灣本島與離島蝶類資源的分類學專著,將分年編纂與出版,中英對照,共計五卷。。2018。臺灣蝶類誌 第一卷 鳳蝶科。行政院農業委員會林務局。239頁。ISBN 9789860551570,GPN 1010602819。
  6. 徐堉峰,黃嘉龍,梁家源。2018。《臺灣蝶類誌第二卷:粉蝶科》內容均依正統分類學方式製作,按科屬種方式編排,每一蝶種介紹,包含中名、異名表列、標本檢查、成蟲形態、世界分佈、在台灣分布、幼蟲寄主植物、生物學及註記;而且每一種類都有雌雄背、腹面標本彩圖、交尾器解剖彩圖,是首部完整論述臺灣本島與離島蝶類資源的分類學專著,將分年編纂與出版,中英對照,共計五卷。
  7. 徐堉峰。2006。台灣蝶圖鑑,第三卷。台灣省立鳳凰谷鳥園。 344 pp。1999台灣蝶圖鑑,第一卷。台灣省立鳳凰谷鳥園。 344 pp。
  8. Hsu, Y. F. & J. A. Powell. Phylogenetic relationships within Heliodinidae and systematics of moths formerly assigned to Heliodines Stainton (Lepidoptera: Yponomeutoidea). 214 pp. University of California Publication in Entomology 124.
  9. 徐堉峰、呂至堅、羅尹廷。 金門常見昆蟲。 內政部營建署金門國家公園 管理處。 191 pp。
  10. 徐堉峰。2002。台灣蝶圖鑑,第二卷。台灣省立鳳凰谷鳥園。383 pp。
  11. 徐堉峰。1999。台灣蝶圖鑑,第一卷。台灣省立鳳凰谷鳥園。 344 pp。

  1. 徐堉峰。2015。絕滅に瀕するソテツ植物を脅かすクロマダラソテツシジミ(The cycad blue, Chilades pandava, a threat to endangered cycad plants)。 Pp. 81-92。矢田 脩主編。熱帶アジアのチョウ(Butterflies of Tropical Asia)。東京,北隆館。
  2. 徐堉峰。2011。台灣水青岡森林的昆蟲。134 -149頁。冰河孑遺的夏綠林 -- 台灣水青岡。行政院農業委員會林務局。臺北市。271頁。
  3. Chiba, H., Y. F. Hsu, & T. Shirozu. 1992. Hesperiidae, pp. 130-132. In Lepidoptera of Taiwan (Heppner, J. B. & H. Inoue, eds). Association for Tropical Lepidoptera. 276 pp.

  1. 王震哲主編(王震哲、林思民、徐堉峰、梁珆碩、梁彧禎、汪仁傑、黃智偉)。師大生態地圖-師大七十回顧叢書。國立臺灣師範大學出版中心。288 頁。
  2. 徐堉峰。2013。臺灣蝴蝶圖鑑 (中):灰蝶。晨星出版社。336頁。
  3. 徐堉峰。2013。徐堉峰。臺灣蝴蝶圖鑑 (下):蛺蝶。晨星出版社。384頁。
  4. 徐堉峰。2013。臺灣蝴蝶圖鑑(上):弄蝶、鳳蝶、粉蝶。晨星出版社。
  5. 楊平世、徐堉峰、侯陶謙著;安英姬繪。鳳翼蝶衣:海峽兩岸鳳蝶工筆彩繪。國立臺灣大學出版中心。159 pp。ISBN:9789860313925
  6. 黃行七、呂晟智、徐堉峰。2010。台灣疑難種蝴蝶辨識手冊。中華民國自然生態保育協會。140頁。
  7. 徐堉峰。2004。近郊蝴蝶。聯經出版社。231 pp。